The dragon keepers and the fledgling dragons are forging a passage up the treacherous Rain Wild River. They are in search of the mythical Elderling city of Kelsingra, and are accompanied by the liveship Tarman, its captain, Leftrin, and a group of hunters who must search the forests for game with which to keep the dragons fed. With them are Alise, who has escaped her cold marriage to the cruel libertine Hest Finbok on order to continue her study of dragons, and Hest's amanuensis, Bingtown dandy, Sedric.